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    Evolutionary theory and mechanism; systematics and phylogenetics; the evolutionary record; evolutionary ecology and behavioural ecology; evolution of human social behaviour.

    All biological phenomena we study today are derived from evolutionary events that occurred in the past. This paper is essential for students majoring in Zoology, but is also of importance to Genetics and Ecology majors and indeed anyone wanting to understand biology in an evolutionary context. In this paper we will explore exactly what evolution is, how it works and what its consequences are. Specific topics covered will range from systematics and phylogenetics, to the evolution of vertebrates and behavioural ecology.

    About this paper

    Paper title Evolutionary Biology
    Subject Zoology
    EFTS 0.15
    Points 18 points
    Teaching period Semester 2 (On campus)
    Domestic Tuition Fees ( NZD ) $1,173.30
    International Tuition Fees Tuition Fees for international students are elsewhere on this website.
    CELS 191 and (BIOL 112 or (HUBS 191 or HUBS 192 or PTWY 131 with at least a B pass))
    Schedule C

    Teaching staff


    Dr Sheena Townsend (ZOOL222 course coordinator)

    Dr Nic Rawlence (ZOOL222 course convenor)

    Dr Travis Ingram

    Teaching fellows:

    Dr Anne Besson

    Paper Structure

    Instruction is delivered via a combination of lectures, assigned readings and labs. Assessment includes project work and writing assignments with both formative and summative components, as well as a final exam.

    Teaching Arrangements

    Two lectures a week and one lab per fortnight. Learning is supported with scheduled tutorials and online learning modules.


    Freeman & Herron, 2014, Evolutionary Analysis (5th ed), Pearson, NJ

    Graduate Attributes Emphasised
    Lifelong learning, Scholarship, Communication, Teamwork.
    View more information about Otago's graduate attributes.
    Learning Outcomes
    Students who successfully complete the paper will develop
    • Knowledge and understanding of the principles of evolution
    • Knowledge of the investigative tools and data used in investigating pattern and process in evolutionary biology


    Semester 2

    Teaching method
    This paper is taught On Campus
    Learning management system


    Stream Days Times Weeks
    A1 Monday 11:00-11:50 29, 42
    Tuesday 09:00-09:50 29-35, 37-42
    Wednesday 09:00-09:50 29-35, 37-42


    Stream Days Times Weeks
    Attend one stream from
    A1 Tuesday 14:00-16:50 29, 31, 33, 35, 38, 40, 42
    A2 Wednesday 14:00-16:50 29, 31, 33, 35, 38, 40, 42


    Stream Days Times Weeks
    A1 Monday 11:00-12:50 32, 34, 39-40
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